Landscape Ecology - Lecture Notes
Some Basic
Concepts in Landscape Ecology
Landscape
Elements
- Landscape elements are spatial units that comprise a
landscape (mosaic). But landscapes can also be conceptualized
as systems with continuously changing gradients.
- Three types of landscape elements are commonly
recognized (The Patch-Corridor-Matrix Model; Forman and Godron
1986 and Forman 1995)
- Patch : "A relatively homogeneous nonlinear
area that differs from its surroundings" (Forman 1995)
- Corridor : A strip of a particular type
that differs from the adjacent land on both sides (as a
conduit, barrier, or habitat)
- Matrix : The background ecosystem or
land-use type in a mosaic, characterized by extensive cover,
high connectivity, and/or major control over dynamics
[Click to zoom in.]
Pattern and Process
- Pattern usually refers to the composition and configuration
of a system.
- Processes refer to actions or activities that create,
modify, or maintain spatial patterns.
- Both pattern and process are scale-dependent.
- "Pattern and process are related to each other" - the
cliche; but is it always true?
- Pattern and process may be interactive, but when?
- Pattern may constrain process, but when?
- Does process always affect pattern?
- They may not be related to each other?
Disturbance
- "Any relatively discrete
event in time that disrupts ecosystem, community, or
population structure
and changes resources, substrate availability, or the physical
environment" (Pickett & White 1985)
Contrast
- The degree of difference among spatial elements, in
particular, among patches, and between patches and underlying
matrix
Spatial
Heterogeneity
- Spatial variation composed of patchiness (discrete changes)
and gradients (gradual changes). Spatial heterogeneity is,
therefore, a more general term than patchiness.
Scale
- The spatial or temporal dimension of an object or process,
characterized by both grain and extent
Grain
- The coarseness in texture of an area, as determined by the
size of patches recognized (i.e., fine grained landscapes
have mostly small patches, while coarse grain has mainly
large patches)
- The finest level of spatial resolution (or the finest
level of temporal resolution) possible with a given data
set; e.g., pixel size for raster data
A comprehensive conceptualization
of scale
(Wu, J. and H. Li. 2006. Concepts of scale and scaling.
Pages 3-15 in J. Wu, K. B. Jones, H. Li, and O. L. Loucks,
editors. Scaling and Uncertainty Analysis in Ecology: Methods
and Applications. Springer, Dordrecht.)
Hierarchy Theory (see
Wu 2013 for details)
- What is a hierarchy or a
hierarchical system?
- What is hierarchy theory?
- How has it been applied in
landscape ecology?
- A simple definition of hierarchy is a partially
ordered set of objects.
- As a theory of complex systems, hierarchy theory was
developed in the framework of general systems theory,
mathematics, and philosophy in the 1960s and 1970s.
- A hierarchically organized system can be seen as a
system in which levels corresponding with progressively
slower behavior are at the top, while those reflecting
successively faster behavior are seen as lower levels in the
hierarchy. Higher levels impose constraints on lower levels,
and thus can be expressed as constants. On the other hand,
the dynamics of the lower levels can be so fast that their
signals are smoothed out at higher levels, and often can be
treated as averages.
- A central property of hierarchy theory in ecology is
that many complex systems have a structure that is
hierarchical and thus "decomposable," implying that both the
analysis and understanding of these systems can be enhanced
by organizing their numerous components into fewer discrete,
interactive units at different levels based on differences
in process rates.
- Process rates (expressed by, for example, cycle time,
response time, or occurrence frequency) are fundamental
characteristics of most systems, and thus may serve as a
common criterion for decomposing. - Other features, such as
tangible boundaries and structural components, also can be
used.
- From the theory, discontinuities in hierarchies for
organizational, spatial, and temporal scales (e.g., breaks
in vegetation pattern, distribution of organism abundance,
distribution of animal body mass, frequency domain of
disturbance) are expected to be a common feature of
ecological systems.
- A main contribution of
hierarchy theory to landscape ecology has been its role in
promoting the study of scale and scaling.
- References:
- Wu, J. 2013. Hierarchy
theory: An overview. Pages 281-302 in: R. Rozzi, S.T.A.
Pickett, C. Palmer, J.J. Armesto, and J.B. Callicott,
(eds). Linking Ecology and Ethics for a Changing World:
Values, Philosophy, and Action. Springer, New York
- Wu, J. 1999. Hierarchy
and scaling. Canadian Journal of Remote Sensing 25(4):
367-380.
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